Lloyd got his start in geology and paleontology at the University of Arkansas, and was introduced to its commercial use through G. H. Cady, doing field studies in southern Illinois for the Mid-Egypt Oil and Gas Company. Lloyd went on to further studies at The University of Kansas and then Yale University, but he continued to do summer commercial work in oil or coal. His studies for the Illinois State Geological Survey Coal Division led to specialization in the fusulinid foraminifera that are common in the limestones and associated shales interbedded with the coal deposits.

Lloyd's first paper on foraminifera described the fusulinellas of the Stonefort Limestone Member of the Tradewater Formation in Illinois. It was published in 1928 in the second volume of the Journal of Paleontology. Recognition of taxonomic problems in this group led to further studies and publication of a paper with Carl O. Dunbar on the fusulinid genera Fusulina, Fusulinella, and Wedekindellina.

Lloyd joined the staff of the U.S. Geological Survey in 1930 and continued his studies of the fusulinids and of upper Paleozoic stratigraphy. His association with Professor Dunbar at Yale continued to yield papers on fusulinid genera, on comparative morphology and evolutionary trends in the fusulinids, and also a monograph on the Pennsylvanian Fusulinidae of Illinois.

Lloyd became interested in obtaining better information on details of the wall structure of the fusulinids and worked with a variety of staining techniques. In a paper on the use of selective stains in paleontology, he showed that many details not normally visible can be brought into sharp contrast through selective staining. One of the most interesting applications was the elaboration of keriothecal structures through staining. His papers on keriothecal wall structure in Fusulina, its significance and resolution by selective stains, and the significance of keriothecal wall structure in Fusulina and its influence on fasuline classification, are examples of this phase of his studies.

Lloyd did considerable work on the smaller foraminifers that are found associated with the fusulinids in some deposits and on the relationship between forams and algae. His outstanding paper on the biology, mineralogy, and diagenesis of some typical late Paleozoic sedentary foraminifers and algal-foraminiferal colonies was published as a special paper of the Cushman Foundation. Diagenesis intrigued Lloyd, not only in the fossils but also in the sediments, and he studied oolitic limestones and fusulinid limestones for evidence of diagenetic and deformational changes. His U.S. Geological Survey Professional Paper on Limestones of Morrow Age in Arkansas and Oklahoma illustrates some of these studies.

Wider interests in paleontology and evolution are well illustrated by his award-winning symposium on the distribution of evolutionary explosions in geologic time, presented at the 1949 meeting of The Paleontological Society.

Lloyd was always interested in helping those who were doing field studies in rocks of late Paleozoic age, and he would visit the mapping parties, collect fusulinids, and give preliminary age assignments for the rocks. Many of these studies resulted either in citations in publications or in cooperative reports with the field geologists. His reports with A. A. Baker on the Wasatch area, Utah, with J. D. Love and N. M. Denson on the Hartville area, Wyoming, and with P. B. King on the Sierra Diablo area, Texas, are examples of these cooperative efforts.

Lloyd was a close friend of Dr. Cushman and his entire family, and the two men worked closely together for many years. The Cushman Foundation is especially aware of the many contributions Lloyd made in getting the Foundation established and in seeing it through its early years. Lloyd guided the group that organized and then incorporated the foundation, and he served as its fourth president. His interest continued as a Director and, now, as Honorary Director.

It is with great pleasure that the Directors of the Cushman Foundation present Lloyd Henbest the Joseph A. Cushman Award for 1985.

RAYMOND C. DOUGLASS, U.S. Geological Survey

from: Journal of Foraminiferal Research, v. 16, no. 3, p. 173, July 1986